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Convert Gallon to ML Please provide values below to convert gallon (US) [gal (US)] to milliliter [mL], or vice versa. A gallon is a unit of volume specifically regarding liquid capacity in both the US customary and imperial systems of measurement. The US gallon is defined as 231 cubic inches (3.785 liters). In contrast, the imperial gallon, which is used in the United Kingdom, Canada, and some Caribbean nations, is defined as 4.54609 liters. In both systems, the gallon is divided into four quarts. A milliliter (symbol: mL) is a unit of volume that is accepted for use in the international system of units (SI). One milliliter is equal to 1 cubic centimeter (cm3), 1/1,000,000 cubic meters (m3), or 1/1000 liters.